I just realized that I selected Edit > Delete All By Type > History before the last save.
I think that broke it.
I just tested that theory by rebinding all my skeletons to my legs, deleted the history on one of them and it broke. Lol, sorry about that, I'm a goofy n00b. So I think I fixed it, I just won't delete all history anymore.
"The first rule in government spending: 'Why build one when you can have two at twice the price?'" - From the movie 'Contact'.